Squamules: up to 5 mm wide, elongate, adnate to imbricate, weakly concave to weakly convex; upper surface: castaneous to dark brown, dull or shiny, epruinose or partly pruinose, sparingly to strongly fissured; margin: concolorous with upper side, straight or up-turned, crenulate to lobed; upper cortex: up to 60-120 μm thick, composed of thick-walled hyphae with more or less angular lumina, not containing crystals of lichen substances, sometimes containing calcium oxalate; medulla: not containing lichen substances or calcium oxalate; lower cortex: of mainly periclinally oriented hyphae, not containing calcium oxalate; lower surface: medium brown, K-; Apothecia: up to 1.5 mm diam., laminal, immarginate or marginate when young and later becoming immarginate, dark brown to black, epruinose or yellow pruinose; ascospores: 10-13 x 5-7 μm; Pycnidia: laminal, immersed; conidia: bacilliform, 6-9 x 1 μm; Spot tests: upper cortex and medulla K-, C-, KC-, P-; Secondary metabolites: none.